Liriodendron tulipifera flower

The University of North Carolina
A Department of the North Carolina Botanical Garden


Collectors of the UNC Herbarium
compiled by Carol Ann McCormick, NCU Curator
Special thanks to
Dr. Jeff Nekola, Department of Biology, University of New Mexico
Alexandra Sharp, Information Services Librarian, Tennessee Wesleyan University
Alan Zahorsky, Reference & Instruction Librarian, University of Mount Union

Ellsworth Bethel
(20 June 1863 – 8 September 1925)


The University of North Carolina Chapel Hill Herbarium (NCU) curates about a dozen fungal specimens collected by Ellsworth Bethel.  It is likely that we will find more as we continue to catalog our collections. 

Ellsworth Bethel’s specimens are distributed widely in herbaria across North America.  He was interested in all aspects of natural history, so collected not only fungi, but also lichens, bryophytes, algae and vascular plants.  According to mycologist Fred Seaver, Bethel was “an omnivorous collector.”1

Among non-scientists, Bethel’s most lasting contribution is the names of several peaks in the Rocky Mountains:
Ellsworth Bethel (1863-1925) was a Denver high school teacher who championed the cause of naming area peaks for Native American tribes. In 1914 he sketched a map of northern Colorado, complete with his recommendation for names of mountains at the time unnamed. He sent it to the U.S. Board on Geographic Names in Washington, D.C.  His main goal was to represent 10 tribes connected with Colorado history by peaks visible from Denver. The board accepted some of his recommendations (such as the one for Mount Acoma) but rejected others. Bethel was angry and disappointed by the board’s decision to throw out some of his ideas, but they accepted enough of his suggestions that his influence still lives today in the mountains of the Indian Peaks Wilderness Area.
Foster, Lisa.  2005.  Rocky Mountain National Park:  The Complete Hiking Guide.  Englewood, Colorado:  Westcliffe Publishers.  Page 315.

Mount Acoma, Grand County, Colorado
40.1761 latitude, -105.7828 longitude
photo by Mike Offerman, 20109

Ellsworth Bethel was born on 20 June, 1863 in Ohio.  His father, William Smith Bethel, was a farmer.  His mother was Phebe Ann Price Bethel, and his sister, Sarah, was two years his junior.  Bethel attended Scio College (now University of Mount Union), though it is unclear for how long.13 Bethel was a Senior Student of Science at Grant Memorial University (now Tennessee Wesleyan University) in 1885-86.14  He was awarded an honorary Masters degree from the University of Denver in 1905.1

According to Fred Seaver, Bethel moved to Colorado in 1890 to teach high school (both the 1900 and 1910 Federal Census list Bethel’s occupation as “school teacher”).   In 1917 he joined the United States Department of Agriculture’s Forest Pathology laboratory and stayed in that job until his death in 1925.1

On October 3, 1924, at age 61 Ellsworth Bethel married 31 year old Dorothy Stokley [or Stokeley].1,7  Ellsworth Bethel died on the 8th of September, 1925 and is buried in Crown Hill Cemetery, Wheat Ridge, Jefferson County, Colorado. 


Taxa described by Bethel:
Puccinia interveniens Bethel (in Blasdale, University of Calif. Publ. Bot. 7:  119.  1919.)
Cronartium occidentale Hedgc., Bethel & N. R. Hunt (J. Agric. Res., Washington 14:  415.  1918.)

Taxa named in Bethel’s honor:
Betheliella Cockerell, 1924 (type species is Betheliella calocharti Cockerell, 1924):  a genus of bees, now considered to be in the genus Dufourea Lepeletier

Asphondylia betheli Cockerell :  prickly pear fruit gall midge – Bethel collected the larvae for Cockerell ( )

Oligotrophus betheli
Felt:  Juniper tip midge, host plant is Rocky Mountain juniper, Juniperus scopulorum

Rhopalomyia betheliana Cockerell:  a midge causing galls on Artemisia (Asteraceae)

Swertia bethelii Harold St. John (Gentianaceae) :  holotype collected by Ellsworth Bethel 17 May 1923 in California, Shasta County, Fall River Mills ; “The new species S. bethelii occurs in the Sierra Mountains along the Pitt River, California.” p. 22 of St. John, Harold.  1941.  Revision of the genus Swertia (Gentianaceae) of the Americas and the reduction of Frasera.  The American Midland Naturalist 26(1):  1-29.  This species is now considered as Frasera albicaulis var. modocensis (H. St. John) N. H. Holmgren

Physarum bethelii T. Macbr. Ex G. Lister (Monogr. Mycetozoa, Edn 2 (London):  57 (1911) – a slime mold

Roestelia bethelii F. Kern (Bull. Torrey Bot. Club 34:  461.  1907.) – a fungus

Aecidium bethelii Arthur (Bull. Torrey Bot. Club 47:  476.  1920.) – a fungus

Uromyces bethelii Arthur (Bull. Torrey Bot. Club 51:  58.  1924.) – a fungus

Peridermium bethelii Hedgc. & Long (Phytopathology 3(4):  251.  1913.) – a fungus

Frasera albicaulis Griseb. var. modocensis (H. St. John) N.H. Holmgren
(synonym:  Swertia bethelii)

photo by Keir Morse, 2009, courtesy of CalFlora10

Seaver, Fred J. 1926.  Ellsworth Bethel.  Mycologia 18(4):  187-188.

Although not exclusively a mycologist [Ellsworth Bethel] was sufficiently prominent in this field to deserve more than a passing mention. Born in Ohio, June 20, 1863, he migrated to Colorado in 1890, where he soon attained prominence as a teacher and in all-around naturalist. He was a graduate of Scio College and of East Tennessee Wesleyan University, and in 1905 he received an honorary degree of M.A. from the University of Denver.

My own personal acquaintance with Mr. Bethel is almost coextensive, in point of time, with my interest in mycology. From about 1904 until the day of his death (September 8, 1925) we were rather closely associated in things mycological and it is difficult to go far in the herbarium without encountering fungi collected in Colorado by Bethel.

When I first visited Colorado in 1910, Mr. Bethel’s company on a number of trips into the mountains was a great help and inspiration. Several days were spent together at Tolland; also in Geneva Creek Canyon and in the vicinity of Boulder.  On all of these occasions he seemed to have unlimited physical energy and boundless enthusiasm for the work. He was an omnivorous collector.

While the number of mycological titles accredited to him is surprisingly small, his collections have furnished groundwork for a number of articles contributed by other mycologists.  In “The Myxomycetes of Colorado. II” Dr. W. C. Sturgis states: “Up to 1907 I had the invaluable assistance of Mr. Bethel, of Denver, and able and tireless collector, thoroughly acquainted with all parts of Colorado.” Of the 94 species of slime-moulds reported from Colorado in the first paper by Dr. Sturgis a very large percent represented the results of the field work of Mr. Bethel in this particular group. One species of slime-mould, Physarum bethelii (Macbr.) Lister, has been dedicated to him and a fine colored illustration published by Miss Lister, the English authority on the slime-moulds. 

He was especially interested in plant rusts, beginning their collection shortly after going to Colorado, and here again contributed much field information for other workers as well as several articles on his own account. One species, Gymnosporangium betheli (Roestelia betheli Kern), has been dedicated to him by Dr. F. D. Kern now of the Pennsylvania State College.  Also Aecidium betheli and Uromyces betheli by Dr. J. C. Arthur and Peridermium betheli by G. G. Hedgcock and W. H. Long, the latter species now regarded as a synonym of Cronartium comandrae.

And in other fields of mycology he showed similar interest. Many interesting specimens of discomycetes have been sent to us in the course of the years. One cup-fungus, Godronia betheli, named for him by the writer, was found to have an earlier name so must be relegated to synonymy. 

In 1917 he began work for the United States Department of Agriculture in the office of Forest Pathology and was connected with this office until the time of his death. On October 3, 1924, he married Miss Dorothy S. Stokeley of Philadelphia.

He has been a subscriber to MYCOLOGIA and a loyal supporter of this periodical from its earliest inception. His loss will be keenly felt by mycologists.


Cockerell, T. D. A.  1926.  Ellsworth Bethel.  Science 63(1625):  201-202. 

ELLSWORTH BETHEL was born at Smyrna, Ohio in 1863. In 1890 he became instructor in biology in the Eastside High School, Denver, a position which he held until 1917. During this long period he enthusiastically studied plant and animal life of Colorado, paying special attention to those obscure and difficult groups, such as the fungi and slime molds, which had been little investigated in the West. Many hundreds of pupils passed through his hands, and to the great majority he was able to communicate a fair measure of his own interest in nature. Some eventually became competent investigators, and all spoke of him with affection and respect. As a collector, Bethel was extraordinarily active, by no means confining himself to those groups which he personally studied. He was continually calling attention to the facts and problems in such fields as entomology and conchology, in which he did not pretend to be an expert. The great accumulation of plants, especially fungi, in the State Museum at Denver, ran far ahead the possibility of critical study with the available resources. It was always Bethel’s hope that he would be able, in his later years, to thoroughly revise many of these materials. Unfortunately, the state of Colorado could not appreciate the importance of work on the native fauna and flora, and in the absence of financial support progress was difficult.  Thus Bethel was led to take up work on economic mycology with the U.S. Department of Agriculture, and in this field has left a distinguished record.

Bethel’s scientific discoveries were numerous. In the Grand Canyon of the Colorado, close to the trail where hundreds of tourists pass every year, his keen eyes detected a peculiar snail, which proved entirely new to science, and was named after him by Professor Junius Henderson.*  In California, one April day in 1923, Bethel picked a mariposa lily, and noted an unusual looking bee in the flower.  It turned out to be a new genus, which received the name Betheliella.  Among insects, those forming galls came in for special attention, and many new species were discovered. There are very few whose natural history interests are so wide, and fewer still who have the sort of intuition which leads them to collect unknown or little-known species in groups they do not personally study. In these respects Bethel combined the qualities of a naturalist of the old school with aptitudes more characteristic of modern times.  He was a pioneer of the West, and although cut short in his prime, and variously handicapped during his life, he left a name which will not be forgotten.

Mr. Haven Metcalf, Bethel’s chief in Washington, sends me the following statement, with permission to quote:

In 1917 Bethel was appointed to the office of Forest Pathology as an associate pathologist, at the time we were in confusion over the question whether the white pine blister rust was or was not a native of America. A rust had been collected in Kansas many years ago, which was passing in the literature as blister rust. Suspicion pointed to the probability that this was a rust having its alternate stage on some pine other than a white pine, and Bethel was the only man to whom we could turn for help on this particularly complicated matter. His work played the dominant part in leading to the demonstrations at the Western rust was a new species, Cronartium occidentale, with its alternate stage on pinon pine, thus disposing of the fear that the white pine blister rust might be native. Upon this conclusion rests the value of the fall campaign against the white pine blister rust. Bethel’s knowledge of the rusts of the Rocky Mountain region, or indeed of the whole West, was prodigious. He was the best collector that I ever saw in action… He is profoundly missed by a large group of coworkers and correspondence. There is no one to take up his work.

Other botanists express themselves in similar terms, all emphasizing Bethel’s tremendous industry and enthusiasm, and readiness to assist others working in the same field. Ellsworth Bethel died suddenly at Denver, Colorado, September 8, 1925.


*In regards to the new species of snail discovered by Ellsworth Bethel, consider this information from Dr. Jeff Nekola of the Department of Biology, The University of New Mexico, Albuquerque:  “The ‘species’ [Bethel’s] obituary was referring to is Sonorella betheli Henderson, 1914.  This turns out to be a total [error].  From Pilsbry (1939):  “Misled by a mistaken locality given by the collector [Ellsworth Bethel], Junius Henderson described rather small specimens (diameter 20.5 to 21.5 mm) as Sonorella betheli.  In 1913 after visiting the Grand Canyon, where the thought that shell was picked up, the late Prof. Ellsworth Bethel spent some time in California.  His chief interest was botanical.  Probably these shells were taken near Los Angeles; the type in the University of Colorado Museum and a cotype, 109733 A.N.S.P., both of which I have seen, agree fully with [Helminthoglypta] traski from that vicinity.”  Nekola adds, “H. traski is the common Helminthoglypta of the Santa Monica mountains, though it extends in to eastern and central parts of the Santa Ynez as well.  There is also Oreohelix haydeni betheli  Pilsbry & Cockerell, 1913.  It’s from Glenwood Springs, Colorado along steep bluffs north of the Grand River.”11  So… in the end, Bethel gets a snail… but a different animal from that the one described in Cockerell’s obituary in Science.


Oreohelix haydeni betheli Pilsbry & Cockerell
photo courtesy of Bagni Liggia, Worldwide Mollusc Species Database


Herbaria which curate Bethel’s specimens include 2,3,4,5,6
ISC      Iowa State University                                      vascular plants, fungi
BRY    Brigham Young University                              vascular plants
CM      Carnegie Mellon Museum of Natural History      vascular plants
COLO  University of Colorado                                    vascular plants, lichens
CS        Colorado State University                                vascular plants
DBG:KHD    Denver Botanical Garden                        vascular plants, fungi
F          Field Museum                                                  vascular plants, fungi, bryophytes
IND     Indiana University                                            vascular plants
USU:UTC     Utah State University                              vascular plants
MIN    J.F. Bell Museum of Natural History                vascular plants, fungi, lichens
MO     Missouri Botanical Garden                                vascular plants
NY     New York Botanical Garden                              vascular plants, fungi, lichens, algae,                                             
NYS   New York State Museum                                   fungi
NCSC North Carolina State University                         vascular plants
OSC:OSU     Oregon State University                           vascular plants, fungi
RM     Rocky Mountain Herbarium                              vascular plants
SUU   Southern Utah University                                   vascular plants
NO     Tulane University                                               vascular plants
ARIZ  University of Arizona                                        vascular plants
UCR   University of California, Riverside                     vascular plants
ILL     University of Illinois                                           vascular plants, fungi
MICH University of Michigan                                       vascular plants, fungi, lichens, algae
RENO University of Nevada                                          vascular plants
WIS    University of Wisconsin, Madison                       vascular plants, fungi
PH      Academy of Natural Science, Drexel University         fungi
BRU   Brown University                                                  fungi, algae
CUP   Cornell University                                                  fungi
FH     Farlow Herbarium of Harvard University               fungi, lichens, bryophytes
PDD   Landcare, New Zealand                                          fungi
LSUM Louisiana State University                                     fungi
MU     Miami University                                                   fungi, algae
MSC   Michigan State University                                      fungi, algae
PUR & PUL Purdue University                                         fungi
CHRB Rutgers University                                                  fungi
BPI     United States National Fungi Collection                  fungi
ARIZ  University of Arizona                                              fungi
UBC   University of British Columbia                                fungi
UC      University of California, Berkeley                           fungi, algae
CINC  University of Cincinnati                                           fungi
FLAS  University of Florida                                                fungi
NEB    University of Nebraska                                            fungi
NCU    University of North Carolina Chapel Hill                fungi
TENN  University of Tennessee, Knoxville                         fungi
VT       University of Vermont                                             fungi
WTU    University of Washington                                        fungi
RMS     University of Wyoming                                           fungi
CFMR  USDA Forest Serv., Center for Forest Mycology Research  fungi
FPF      USDA Forest Serv., Rocky Mt. Research Station   fungi
WSP     Washington State University                                    fungi
US        United States National Herbarium                            lichens, bryophytes
YU       Yale University                                                        algae, bryophytes




Bethel, E. 1899.  Notes and News.  The Plant World 2(9):  154.
-----.  1899.  A new method of drying botanical driers.  The Plant World 2(12): 199-200.
Bethel, E. 1911.  Notes on some Gymnosporangium in Colorado.  Mycologia 3(3):  156-160.
-----.  1917.  Puccinia subnitens and its aerial hosts.  Phytopathology 7:  92-94.
-----.  1919.  Puccinia subnitens and its aerial hosts, II.  Phytopathology 9: 193-201.
Hedgcock, G. G., E. Bethel, and N. R. Hunt.  1918.  Pinon blister rust.  J. of Agricultural Research 14(10):  411-424.



1.  Seaver, Fred J. 1926.  Ellsworth Bethel.  Mycologia 18(4):  187-188.
2. searched on 29 August 2017.
3. searched on 29 August 2017.
4. searched on 29 August 2017.
5. searched on 29 August 2017.
6. searched on 29 August 2017.
7.  Colorado, County Marriages and State Indexes, 18622006 [database on-line].  Lehi, UT, USA: Operations, Inc., 2016.  Original data:  Marriage Records.  Colorado Marriages.  FamilySearch, Salt Lake City, UT. 
8.  Cockerell, T. D. A.  1926.  Ellsworth Bethel.  Science, New Series 63(1625):  201-202.
9.  Acoma, Mount.  accessed on 29 August 2017.
10.  Frasera alibcaulis Griseb. var. modocensis (H. St. John) N. H. Holmgren.  accessed on 29 August 2017.
11.  Pers. comm. Email Nekola to McCormick 29 August 2017.
12.  Oreohelix.”  WMSDB – Worldwide Mollusc Species Data Base by Bagni Liggia, Genova, Italy.  accessed on 30 August 2017.

13.  Pers. comm.  Email Alan Zahorsky, University of Mount Union, to McCormick 29 August 2017.
14.  Pers. comm.  Email Alexandra Sharp, Tennessee Wesleyan University, to McCormick 30 August 2017.


   Curriculum in Ecology                 North Carolina Botanical Garden               Biology Department
      Curriculum                               North Carolina                                 UNC

         In Ecology                              Botanical Garden                   Biology Department


University of North Carolina Chapel Hill Herbarium (NCU)
CB# 3280, Coker Hall
120 South Road
University of North Carolina at Chapel Hill
Chapel Hill, NC 27599-3280
phone: (919) 962-6931


Last Updated: 29 August  2017