Liriodendron tulipifera flower

The University of North Carolina
A Department of the North Carolina Botanical Garden


Collectors of the UNC Herbarium
Compiled by Carol Ann McCormick, Asst. Curator, NCU

Donald E. Stone
(10 December 1930 Ė 4 March 2011)


The University of North Carolina Herbarium (NCU) has catalogued about a dozen specimens collected by Don Stone.However, in the course of his research, he frequently consulted NCUís specimens, and hundreds of specimens of Carya and Juglans in our collection were annotated by him.



Don Stone.
Image courtesy of Duke University
Biology Department website

Anon. 2011.Obituary:Donald E. Stone 1930-2011.Flora of North America Newsletter 25(1):15.

Donald Eugene Stone died from a short bout with cancer on Friday, March 4, 2011, in Durham, North Carolina.Don was born on December 10, 1930, and grew up in Eureka, California.Don did his undergraduate work at Humboldt State College and the University of California at Berkeley.He remained at Berkeley and was awarded his Ph.D. in Botany in 1957.Subsequently, he taught at Tulane University for six years.In 1963, he joined the Botany Department at Duke University and taught at Duke for the remainder of his career.In 1969-70, Don took a one-year leave of absence to serve as the Associate Program Director in Systematic Biology at NSF [National Science Foundation].

In 1976, while teaching full-time at Duke, he began shepherding the Organization for Tropical Studies (OTS) as its Executive Director, expanding the consortium from 20 to more than 60 universities, colleges, museums, and research institutions.During this period he enhanced the Organization's field-based graduate courses and created an on-the-ground training program for policy makers.Most importantly, he strengthened OTS's three biological field stations in Costa Rica, La Selva, Las Cruces, and Palo Verde, as major research centers, and, in particular, established the La Selva station as one of the most important sites in the world for research in tropical biology.In the early 1980s, under Don's guidance, OTS took a leadership role, along with the John D. and Catherine T. MacArthur Foundation, the World Wildlife Fund, and The Nature Conservancy, in establishing a protected, 47,000-hectare, forested corridor from the Braulio Carrillo National Park, located in the central highlands of Costa Rica, to La Selva, more than 35 miles away in the Caribbean lowlands.As s result of these efforts, in 1985 OTS was the first organization to be awarded the John and Alice Tyler Prize for Environmental Achievement.

When Don retired from OTS in 1996, he served as the chair of the Botany Department at Duke for three years.In 2000, he joined the OTS Board of Visitors, which he formed in 1992, and from 2003 to 2005 he served in a volunteer capacity as OTS's Interim Executive Director during an 18-month search for the current CEO.

Don's own research interest centered on the systematics and evolution of temperate and tropical plants using biochemistry, cytotaxonomy, comparative anatomy, and comparative morphology in the walnut family (Juglandaceae), and pollen development in the ginger family (Zingiberales).In addition to many other publications, he contributed Juglandaceae to the floras of Mesoamerica, Nicaragua, Costa Rica, and China as well as Juglans and Carya to the Flora of North America, Volume 3.Memorial contributions may be sent to OTS, Box 90633, Durham, NC27708-0633.


Don Stone is survived by his spouse, Beverly Stone.


Stone, Donald E. (1961)Ploidal level and stomatal size in the American hickories.Brittonia 13(3):293-302.

----- (1962)Affinities of a Mexican endemic, Carya palmeri, with American and Asian hickories.American Journal of Botany 49(3):199-212.

----- (1963)Pollen size in hickories (Carya).Brittonia 15(3):208-214.

----- (1964)New chromosome counts for two species of hickory (Carya). Brittonia16 (2):230.

Stone, Donald E., George A. Adrouny and Salpi Adrouny (1965)Morphological and chemical evidence on the hybrid nature of bitter pecan, Carya x Lecontei.Brittonia 17(2):97-106.

Stone, Donald E. (1968)New World Juglandaceae:A new species of Alfaroa from Mexico.American Journal of Botany 55(4):477-484.

Stone, Donald E., George A. Adrouny and Robert H. Flake (1969)New World Juglandaceae, II.Hickory nut oils, phonetic similarities, and evolutionary implications in the genus Carya.Am. J. Bot. 56(8):928-935.

Stone, Donald E. (1970)Evolution of cotyledonary and nodal vasculature in the Juglandaceae.Am. J. Bot. 57(10):1219-1225.

----- (1972)New World Juglandaceae, III.A new perspective of the tropical members with winged fruits.Annals of the Missouri Botanical Garden 59(2):297-322.

----- (1973)Patterns in the Evolution of Amentiferous Fruits.Brittonia25 (4): ††371-384.

Kress, W. John, Donald E. Stone and Susan C. Sellers (1978)Ultrastructure of exine-less pollen:Heliconia (Heliconiaceae).Am. J. Botany 65(10):1064-1076.

Stone, Donald E, Susan C. Sellers and W. John Kress (1979)Ontogeny of exineless pollen in Heliconia, a banana relative.Annals of the Missouri Botanical Garden 66(4):701-730.

----- (1981)Ontogenetic and evolutionary implications of a neotenous exine in Tapeinochilos (Zingiberales:Costaceae) pollen.Am. J. Botany 68(1):49-63.

Kress, W. John and Donald E. Stone (1983)Morphology and phylogenetic significance of exine-less pollen of Heliconia (Heliconiaceae).Systematic Botany 8(2):149-167.

James W. Hardin and Donald E. Stone (1984)Atlas of foliar surface features in woody plants, VI.Carya (Juglandaceae) of North America.Brittonia 36(2):140-153.

Kress, W. John and Donald E. Stone (1993)Morphology and flora biology of Phenakospermum (Strelitziaceae), an arborescent herb of the Neotropics.Biotropica 25(3):290-300.

Manos, Paul S. and Donald E. Stone (2001)Evolution, phylogeny, and systematics of the Juglandaceae.Annals of the Missouri Botanical Garden 88(2):231-269.

Manos, P.S., P.S. Solits, D.E. Soltis, S.R. Mancheser, S.-H. Oh, C.D. Bell, D. L. Dilcher and D. E. Stone (2007)Phylogeny of extant and extinct Juglandaceae inferred from the integration of molecular and morphological data sets.Systemtic Biology 56:1-19.


†† Curriculum in Ecology†††††††††††††††††North Carolina Botanical Garden†††††††††††††††Biology Department
†††††††Curriculum†††††††††††††††††††††††††††††† North Carolina†††††††††††††††††††††††††††††††† UNC

††††††††††In Ecology†††††††††††††††† †††††††††††††Botanical Garden†††††††††††††††††† Biology Department


University of North Carolina Herbarium
CB# 3280, Coker Hall
University of North Carolina
Chapel Hill, NC 27599-3280
phone: (919) 962-6931
fax: (919) 962-6930

Last Updated: 28 October 2012